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Sackler Faculty Members

The Sackler School of Graduate Biomedical Sciences faculty is comprised of about 150 members. Our faculty members are listed below, and we invite you to find out more about our faculty and their research by clicking on their names. Each faculty member's page includes a short research description, her/his program affiliation and, in many cases, a link to the faculty member's more detailed web pages.

Please note: Faculty members who have an asterisk before their names are no longer accepting students for Ph.D. thesis research; however, they are still active in graduate education.

   
 
A B C D E F G H I J K L M N O P Q R S T U V W X Y Z

A

Acacia Alcivar-Warren, D.V.M., Ph.D. Associate Professor of Environmental and Population Health, School of Veterinary Medicine
Genetics 
Molecular and developmental genetics.

David A. Adler, M.D.  Professor of Psychiatry
Clinical Research
Mental Health Services Research

* Mohammed S. Anwer, D.M.V.H., Ph.D. Professor of Pharmacology and Experimental Therapeutics
Pharmacology & Experimental Therapeutics 
Hepatotoxicity; pathophysiology of cholestasis.

* Irwin M. Arias, M.D. Professor of Physiology
Cellular & Molecular Physiology 
Normal and abnormal transport of bile acids & pigments; pathobiology of liver injury and multidrug resistance.



B

William W. Bachovchin, Ph.D. Professor of Biochemistry
Biochemistry 
Serine proteinases: structure, mechanism and biological functions; NMR spectroscopy; design of synthetic agents to manipulate T cells.

Vaidyanathapuram S. Balakrishnan, M.D., M.R.C.P. Associate Professor of Medicine
Clinical Research 
Chronic kidney disease, dialysis, transplantation, kidney stones

James D. Baleja, Ph.D. Associate Professor of Biochemistry
Biochemistry 
Molecular recognition; structure determination using NMR; structure-based drug design.

Debbie Beasley, Ph.D. Associate Professor of Medicine
Cellular and Molecular Physiology 
Role of innate immunity and smooth muscle cells in early atherogenesis

Martin Beinborn, M.D. Assistant Professor of Pharmacology and Experimental Therapeutics
Pharmacology & Experimental Therapeutics 
Molecular processes that lead to ligand-indcued activation of peptide hormone receptors.

Margery C. Beinfeld, Ph.D. Professor of Pharmacology and Experimental Therapeutics
Pharmacology & Experimental Therapeutics 
Neuropeptide biosynthesis, processing, and release.

Evan M. Benjamin, M.D. Associate Professor of Medicine
Clinical Research
Quality of care and patient safety

Jamshed Bharucha Ph.D. Professor of Psychology, School of Arts and Sciences
Neuroscience
Cognitive and neural basis of the perception of music.

Diana W. Bianchi, M.D. Natalie V. Zucker Professor of Pediatrics, Obstetrics and Gynecology
Genetics 
Human fetal genetics; genetic testing; fetal gene therapy.

Robert Blaustein, M.D., Ph.D. Assistant Professor of Medicine
Neuroscience  Biochemistry
Relationship between structure and function in potassium channels

* Eunice I. Bloomquist, Ph.D. Professor of Physiology
Cellular & Molecular Physiology 
Smooth muscle; airway inflammation.

A. Andrew Bohm, Ph.D. Associate Professor of Biochemistry
Biochemistry 
Cell Signal Transduction Mechanisms; Protein Structure Analysis by X-Ray;Crystallography

Peter H. Brodeur, Ph.D. Associate Professor of Pathology
Genetics  Immunology 
Immunoglobulin genetics; molecular organization of variable region genes; expression of V genes and V gene families; development of the antibody repertoire.

Rachel J. Buchsbaum, M.D. Assistant Professor of Medicine
Genetics
Molecular mechanisms of metastasis.

Peter A. Bullock, Ph.D. Professor of Biochemistry
Biochemistry  Genetics 
DNA replication in eukaryotic cells; mechanism of genetic recombination.

Stephen S. Bunnell, Ph.D. Assistant Professor of Pathology
Immunology 
Signaling complexes induced by the T cell receptor.



C

Andrew Camilli, Ph.D. Professor of Molecular Biology and Microbiology
Molecular Microbiology 
Host-pathogen interactions using genetic and molecular biological approaches.

John J. Castellot, Jr., Ph.D. Professor of Anatomy and Cellular Biology
Director of the Cell, Molecular, and Developmental Biology Program
Cell, Molecular & Developmental Biology  Pharmacology & Experimental Therapeutics 
Cellular and molecular mechanisms of regulation of vascular cell proliferation.

Al Charest, M.Sc., Ph.D. Assistant Professor of Neurosurgery
Neuroscience
Brain cancer and pre-clinical applications in genetically engineered mouse models.

Jinkun (Jake) Chen, D.D.S., Ph.D. Professor of General Medicine, School of Dental Medicine
Cell, Molecular & Developmental Biology
Function and control of expression of bone sialoprotein (BSP) in bone and other mineralized tissue. 

Brent H. Cochran, Ph.D. Professor of Physiology
Cellular & Molecular Physiology 
Cell growth regulation.

John M. Coffin, Ph.D. Professor of Molecular Biology and Microbiology
Genetics  Molecular Microbiology 
Molecular biology of retroviruses.

Joshua T. Cohen, Ph.D. Research Associate Professor of Medicine
Clinical Research 
Applying decisions analytical techniques to the analysis of medical interventions and environmental problems.

Michael H. Court, B.V.Sc., Ph.D. Assistant Professor of Pharmacology and Experimental Therapeutics
Pharmacology & Experimental Therapeutics, Genetics  
Drug metabolism, specifically genetic deficiencies in various animal species with particular emphasis on the UDP-glucuronosyl transferases.

Lidija Covic, Ph.D. Assistant Professor of Medicine
Biochemistry
Role of angiogenesis factors in PAR-dependent tumor cell biology

* Janet M. Cowan, Ph.D. Associate Professor of Pediatrics
Genetics 
Cytogenetic analysis of tumors.

Daniel H. Cox, Ph.D. Associate Professor of Neuroscience
Neuroscience  Biochemistry
Relationship between the structures of potassium channel proteins and their function.



D

* David A. Damassa, Ph.D. Professor of Anatomy and Cellular Biology
Cell, Molecular & Developmental Biology 
Reproductive neuroendocrinology.

Olaf Dammann, M.D. Professor of Pediatrics
Clinical Research 
Perinatal and developmental neuroepidemiology

Jonathan M. Davis, M.D. Professor of Pediatrics
Clinical Research  Pharmacology and Experimental Therapeutics
Oxidant/antioxidant balance in the pathogenesis of various disease processes of newborns

Alexei Degterev, Ph.D. Assistant Professor of Biochemistry
Biochemistry
Regulation of cell death.

* James F. Dice, Jr., Ph.D. Professor of Physiology
Cellular & Molecular Physiology 
Intracellular protein degradation and cellular mechanism of aging

* James W. Drysdale, Ph.D. Professor Emeritus of Biochemistry
Biochemistry 
Identification of the hemochromatosis gene.

Thomas Ducibella, Ph.D. Professor of Obstetrics and Gynecology
Cell, Molecular & Developmental Biology 
Mammalian oocyte maturation and in vitro fertilization.

Kathleen Dunlap, Ph.D. Professor of Neuroscience and Director of the Neuroscience Program
Cellular & Molecular Physiology  Neuroscience 
Modulation and physiological function of voltage-dependent channels.



E

* Al Walid I. El-Bermani, M.D., Ph.D. Associate Professor of Anatomy and Cellular Biology
Cell, Molecular & Developmental Biology 
Lung innervation; vascular response to leukotriene.



F

Jerry R. Faust, Ph.D. Associate Professor of Physiology
Cellular & Molecular Physiology 
Neuronal ceroid lipofuscinosis; isopentenyladenosine in tRNA's; cholesterol homeostasis.

Larry A. Feig, Ph.D. Professor of Biochemistry
Director of the Biochemistry Program
Biochemistry  Genetics  Neuroscience 
Signal transduction pathways mediated by RAS and related GTPases; function of RAS in neuronal calcium signaling.

Michael D. Forgac, Ph.D. Professor of Physiology
Biochemistry, Cellular & Molecular Physiology 
Mechanism of regulation of vacuolar acidification.

Eric Frank, Ph.D. Professor of Physiology
Cellular & Molecular Physiology, Neuroscience 
Molecular mechanisms specifying the development of synaptic connections between neurons

Catherine Freudenreich, Ph.D. Associate Professor of Biology, School of Arts and Sciences
Genetics 
DNA mutation and chromosome fragility in yeast.



G

Jonas Galper, M.D., Ph.D. Professor of Medicine
Cellular & Molecular Physiology
Sterol regulation of cardiovascular disease.

Jonathan Garlick, D.D.S., Ph.D. Professor of Oral and Maxillofacial Pathology, School of Dental Medicine
Cell, Molecular & Developmental Biology
The molecular and cellular mechanisms of cell-cell and cell-stromal interactions in squamous cell cancer.

Grace B. Gill, Ph.D. Associate Professor of Anatomy and Cellular Biology
Cell, Molecular & Developmental Biology  Genetics
Regulation of cell growth and differentiation by post-translational modification of transcription factors

Edward B. Goldberg, Ph.D. Professor of Molecular Biology and Microbiology
Molecular Microbiology 
Genetic design of proteins for biometric, functional, self-assembling nanomaterials.

Robert J. Goldberg, Ph.D. Adjunct Professor of Medicine
Clinical Research 
Coronary heart disease; incidence and survival rates of acute myocardial infarction and out-of-hospital coronary deaths.

Alice B. Gottlieb, M.D., Ph.D. Professor of Dermatology
Clinical Research  Immunology
Treatments for psoriasis and psoriatic arthritis.

David J. Greenblatt, M.D.  Louis Lasagna Professor of Pharmacology and Experimental Therapeutics
Biochemistry  Pharmacology & Experimental Therapeutics 
Clinical psychpharmacology; pharmacokinetics and metabolism.

John Griffith, Ph.D. Associate Professor of Medicine,
Director, Biostatistics Research Center

Clinical Research
Development and application of new statistical methodologies for clinical research, and statistical and analytic support for clinical researchers.  

* Alvar W. Gustafson, Ph.D. Associate Professor of Anatomy and Cellular Biology
Cell, Molecular & Developmental Biology 
Male reproductive system.



H

Akiko Hata, Ph.D. Associate Professor of Biochemistry
Biochemistry, Cell, Molecular and Developmental Biology 
Signaling pathways in vertebrate development.

Victor Hatini, Ph.D. Assistant Professor of Anatomy and Cellular Biology
Cell, Molecular and Developmental Biology, Genetics
Signaling, patterning, morphogenesis and organ growth

Catherine Hayes, D.M.D., D.M.S. Professor of Public Health and Community Service, School of Dental Medicine
Clinical Research
Oral health disparities

Ekaterina Heldwein, Ph.D. Assistant Professor of Molecular Biology and Microbiology
Molecular Microbiology
Structural and mechanistic studies of herpesvirus entry into host cells

Ira M. Herman, Ph.D. Professor of Physiology
Cell, Molecular & Developmental Biology  Cellular & Molecular Physiology 
Molecular control of cell growth, motility, and differentiation.

Richard C. Hermann, M.D., M.S. Associate Professor of Medicine and Psychiatry
Clinical Research 
Methods of quality assessment and improvement in health care.

Patricia L. Hibberd, M.D., Ph.D. Professor of Medicine
Pharmacology and Experimental Therapeutics
International clinical trials of novel anti-infective agents.

Philip Hinds, Ph.D. Professor of Radiation Oncology
Genetics  Biochemistry
G1 cell cycle control and its dysregulation in cancer cells.

Linden Hu, M.D. Associate Professor of Medicine
Immunology  Molecular Microbiology
Host - Pathogen Interaction in Borrelia Infections

Brigitte T. Huber, Ph.D. Professor of Pathology
Genetics  Immunology 
Molecular mechanism of T-cell activation and differentiation; functional and structural analysis of virally encoded superantigens.

Gordon Huggins, M.D.  Assistant Professor of Medicine
Cell, Molecular & Developmental Biology  Genetics
Genetic regulation of heart muscle stem cell development



I

Thereza Imanishi-Kari, Ph.D. Associate Professor of Pathology
Genetics  Immunology 
The study of cellular interactions involved in the creation of the antibody repertoire: its genetic basis and selection in ontogeny.

Ralph Isberg, Ph.D. Professor of Molecular Biology and Microbiology
Molecular Microbiology  Genetics
Intracellular pathogens and host cell membrane trafficking.



J

F. Rob Jackson, Ph.D. Professor of Neuroscience
Genetics  Neuroscience 
Genetic analysis of the nervous system and behavior.

Michele H. Jacob, Ph.D. Associate Professor of Neuroscience
Cell, Molecular & Developmental Biology  Neuroscience  Cellular and Molecular Physiology
Neuronal synapse formation

* Stanley Jacobson, Ph.D. Professor of Anatomy and Cellular Biology
Cell, Molecular & Developmental Biology 
Neurocircuitry and CNS.

Daniel G. Jay, Ph.D. Professor of Physiology
Cellular & Molecular Physiology  Neuroscience 
Molecular mechanisms of axon guidance and nerve growth cone motility.

Douglas M. Jefferson, Ph.D. Associate Professor of Physiology
Cellular & Molecular Physiology 
Genetic diseases that affect liver function.

Peter Juo, Ph.D. Assistant Professor of Physiology
Cellular & Molecular Physiology  Neuroscience
Cell biology of the synapse; localization and regulation of synaptic proteins.



K

David Kaplan, Ph.D. Professor of Biomedical Engineering, School of Engineering
Cell, Molecular & Developmental Biology
The interface between biology and materials science and engineering.

Richard Karas, M.D., Ph.D.  Professor of Medicine
Cell, Molecular & Developmental Biology 
The effect of estrogen on the cardiovascular system.

* John S. Kauer, Ph.D. Professor of Neuroscience
Cell, Molecular & Developmental Biology  Neuroscience 
Analysis of neuronal circuitry.

David Kent, M.D. Associate Professor of Medicine
Clinical Research
Application of multivariate predictive models in cardiac disease & stroke

* Roy L. Kisliuk, Ph.D. Professor Emeritus of Biochemistry
Biochemistry  Pharmacology & Experimental Therapeutics 
Folate enzymes, coenzymes, and antimetabolites.

Alan S. Kopin, M.D. Professor of Medicine
Genetics  Neuroscience  Pharmacology & Experimental Therapeutics 
Molecular analysis of G-protein coupled receptor function in vitro and in vivo.

* Norman I. Krinsky, Ph.D. Professor Emeritus of Biochemistry
Biochemistry 
Metabolism of carotenoids to retinoids and retinoic acid. Role of carotenoids in human vision. Function of antioxidants.

Athan Kuliopulos, M.D., Ph.D. Associate Professor of Medicine
Biochemistry  Genetics 
Signal transduction of peptide receptors; genetic, biochemical, and NMR structural characterization of the thrombin receptor.

Carol A. Kumamoto, Ph.D. Professor of Molecular Biology and Microbiology
Genetics  Molecular Microbiology 
Invasive growth of the fungal pathogen, Candida albicans; mechanisms of antifungal resistance in C. albicans.

Rajendra Kumar-Singh, Ph.D. Associate Professor of Ophthalmology
Genetics  Cell, Molecular & Developmental Biology  Neuroscience
Development and application of gene therapy vectors for retinal diseases.

Charlotte Kuperwasser, Ph.D. Assistant Professor of Anatomy and Cellular Biology
Cell, Molecular & Developmental Biology, Genetics
Matrix and tissue regulation of the molecular pathways of cancer progression

John Kyriakis, Ph.D. Professor of Medicine
Biochemistry
Signal transduction mechanisms recruited by mitogenic, environmental and inflammatory stresses



L

Joseph Lau, M.D. Professor of Medicine
Clinical Research  
Evidence-based medicine and meta-analysis

* Paul C. Leavis, Ph.D. Associate Professor of Physiology
Cellular & Molecular Physiology 
Fetal peptides as anti-cancer agents.

Ronald M. Lechan, M.D., Ph.D. Professor of Medicine
Neuroscience 
Anatomical connectivity and neuroendocrine regulation of the hypothalamus and pituitary.

Janis Lem, Ph.D. Research Associate Professor of Medicine
Cell, Molecular and Developmental Biology  Genetics  Neuroscience
Molecular analysis of phototransduction mechanisms.

Debra J. Lerner, MS, Ph.D. Associate Professor of Medicine
Clinical Research 
Measurement of work disability and health related quality of life for chronically ill adults.

Laurel K. Leslie, M.D., M.P.H. Associate Professor of Medicine
Clinical Research
Identification and treatment of children and adolescent mental health issues.

Andrew Levey, M.D. Professor of Medicine
Clinical Research 
Role of diet and blood pressure in the progression of diabetic and non-diabetic renal disease.

Peter K. Lindenauer, M.D., M.Sc., FACP Associate Professor of Medicine
Clinical Research
Patient safety, clinical practice guidelines, pulmonary disease

Thomas F. Linsenmayer, Ph.D. Professor of Anatomy and Cellular Biology
Cell, Molecular & Developmental Biology 
Extracellular matrices.

Laura Liscum, Ph.D. Professor of Physiology
Director of the Cellular and Molecular Physiology Program
Cellular & Molecular PhysiologyNeuroscience
Intracellular cholesterol transport and metabolism.



M

Michael H. Malamy, Ph.D. Professor of Molecular Biology and Microbiology
Genetics  Molecular Microbiology 
Molecular biology of plasmids & pathogens.

Tim McAlindon, M.D., M.P.H. Associate Professor of Medicine
Clinical Research
Epidemiologic and clinical studies of rheumatic disease; Informatics.

Mitch McVey, Ph.D. Assistant Professor of Biology, School of Arts and Sciences
Genetics
Molecular mechanisms of DNA repair in Drosophila melanogaster.

Joan C. Mecsas, Ph.D. Associate Professor of Molecular Biology and Microbiology
Molecular Microbiology  Immunology
Pathogenic mechanisms of enteric bacteria and the role of Yersinia yops.

Karina F. Meiri, Ph.D. Professor of Anatomy and Cellular Biology
Cell, Molecular & Developmental Biology  Neuroscience  Pharmacology and Experimental Therapeutics
Regulation of neuritogenesis, neuron differentiation and synapse formation.

Michael E. Mendelsohn, M.D. Professor of Medicine
Biochemistry  Cellular & Molecular Physiology   Cell, Molecular & Developmental Biology
Molecular signaling mechanisms in vascular cells; molecular mechanisms of vascular cells; molecular mechanisms of nitric oxide/cyclic GMP action.

Vandana Menon, M.D., Ph.D., M.P.H. Assistant Professor of Medicine
Clinical Research
Novel cardiovascular risk factors in chronic kidney disease.

Simin N. Meydani, D.V.M., Ph.D. Professor of Nutrition, Friedman School of Nutrition Science & Policy
Immunology 
Nutrition, antioxidant, & the immune response; age-related changes in immunity.

Klaus A. Miczek, Ph.D. Professor of Psychology, School of Arts and Sciences
Genetics
Neurobiology and psychopharmacology of social and aggressive behavior; behavioral neuroscience.

Sergei M. Mirkin, Ph.D. Professor of Biology, School of Arts and Sciences
Genetics 
Genome structure and functioning in norm and disease.

Claire L. Moore, Ph.D. Professor of Molecular Biology and Microbiology
Biochemistry  Genetics  Molecular Microbiology 
Post-transcriptional processing of eukaryotic messenger RNA and its role in the regulation of gene expression.



N

Bradford J. Navia, M.D., Ph.D. Associate Professor of Neurology
Genetics 
Genetics of neurodevelopmental disorders; intracellular signals that contribute to HIV-induced neuronal apoptosis.

Heber Nielsen, M.D. Professor of Pediatrics
Cell, Molecular & Developmental Biology
Normal and abnormal regulation of fetal lung development.

Peter J. Neumann, Sc.D. Professor of Medicine
Clinical Research
Cost-effectiveness analysis in health care decision-making.

Dmitry I. Nurminsky Ph.D. Assistant Professor of Anatomy and Cellular Biology
Cell, Molecular & Developmental Biology
Protein – DNA interaction that regulate gene expression



O

Jose M. Ordovas, Ph.D. Professor of Nutrition, Friedman School of Nutrition Science & Policy
Genetics 
Genetics of cardiovascular disease and gene by diet interactions.

* Nanette Roberts Orme-Johnson, Ph.D. Associate Professor Emerita of Biochemistry
Biochemistry  Pharmacology & Experimental Therapeutics 
Subcellular mechanisms of control of steroid hormone synthesis; effects of xenobiotics on steriod hormone synthesis.



P

Noorjahan Panjwani, Ph.D. Professor of Ophthalmology
Biochemistry  Cell, Molecular & Developmental Biology 
Mechanisms of corneal infection and corneal wound healing.

Susan K. Parsons, M.D., M.R.P. Associate Professor of Medicine
Clinical Research  
Pediatric hematology/oncology and health services research.

K. Eric Paulson, Ph.D. Assistant Professor of Radiation Oncology
Biochemistry 
Molecular and cellular regulation of liver function.

Brian J.G. Pereira, M.D.  Professor of Medicine
Clinical Research 
Hepatitis in dialysis and transplantation; cytokines in dialysis, biocompatibility of dialysis membranes and clinical outcomes.

Mercio A. PereiraPerrin, M.D., Ph.D. Professor of Pathology
Immunology 
Roles of surface glycoproteins of trypanosomes and Leishmaniae in virulence; signal transduction in microbial invasion.

Inga Peter, Ph.D.  Assistant Professor of Medicine
Clinical Research
Gene identification in complex diseases

* Andrew G. Plaut, M.D. Professor of Medicine
Immunology 
Mucosal immunity and bacterial infection, IgA proteases and disease

Alexander Poltorak, Ph.D. Assistant Professor of Pathology
Genetics  Immunology  Molecular Microbiology
Use of mouse models to dissect the genetics of innate immunity

Emmanuel N. Pothos, Ph.D. Assistant Professor of Pharmacology and Experimental Therapeutics
Pharmacology and Experimental Therapeutics, Neuroscience
The basic mechanism of synaptic plasticity in the midbrain

 



Q

Henry W. Querfurth, M.D., Ph.D. Associate Professor of Neurology
Neuroscience
Study of muscle pathogenesis as related to Alzheimer's disease protein aggregation.




R

Maribel Rios, Ph.D. Assistant Professor of Neuroscience
Cellular and Molecular Physiology  Neuroscience
Genetic analysis of neurotrophic functions; regulation of affective behavior, food intake and metabolic function by brain derived neurotrophic factor.

Suzanne K. Roffler-Tarlov, Ph.D. Professor of Neuroscience
Cell, Molecular & Developmental Biology  Genetics  Neuroscience 
The catecholamine systems; neurodegeneration.

Irwin H. Rosenberg, M.D. Professor of Physiology, Friedman School of Nutrition Science & Policy
Cellular & Molecular Physiology 
Membrane transport and metabolism of vitamins; calcium bioavailability.

*Naomi Rosenberg, Ph.D. Dean of Sackler School and Professor of Pathology
Genetics  Immunology  Molecular Microbiology 
Interaction and development of malignancy, using an Abelson murine leukemia virus model system.

Michael Rothberg, M.D., M.P.H. Assistant Professor of Medicine
Clinical Research
Diagnostic and treatment dilemmas in general medicine, quality of care.

Michael Rosenblatt, M.D. Dean of the School of Medicine and Professor of Physiology
Cellular and Molecular Physiology  Genetics
Hormone-receptor interactions; mechanisms of metastasis to bone

Ananda L. Roy, Ph.D. Associate Professor of Pathology
Genetics  Immunology 
Molecular mechanisms of tissue-specific transcription; signal transduction in lymphoid cells.

Beverly S. Rubin, Ph.D. Associate Professor of Anatomy and Cellular Biology
Cell, Molecular & Developmental Biology 
Neuroendocrine control of gonadtropin secretion and reproductive behaviors.



S

Dana Safran, ScD Associate Professor of Medicine
Clinical Research 
Assessment of primary care performance under different models of health care delivery.

G. Gary Sahagian, Ph.D. Professor of Physiology
Cellular & Molecular Physiology 
Role of lysosomal trafficking in normal and diseased processes.

* Satyapriya Sarkar, Ph.D. Professor Emeritus of Anatomy and Cellular Biology
Cell, Molecular & Developmental Biology  Genetics 
Structure, function, and evolution of vertebrate muscle genes; mechanism of Ca2+ regulation of muscle contraction.

Mark J. Sarnak, M.D., M.S. Associate Professor of Medicine
Clinical Research
Cardiovascular disease in chronic kidney disease.


Brian S. Schaffhausen, Ph.D. Professor of Biochemistry
Biochemistry  Genetics  Molecular Microbiology 
Mechanism of neoplastic transformation; genetic and biochemical analysis of protein modifications; signal transduction.

Christopher H. Schmid, Ph.D. Professor of Medicine
Clinical Research 
Development of statistical analytic methods in clinical research with an emphasis on Bayesian techniques and meta-analysis.

Gavin R. Schnitzler, Ph.D. Assistant Professor of Biochemistry
Biochemistry  Genetics 
Function of human chromatin remodeling complexes in transcriptional control.

James E. Schwob, M.D., Ph.D. Professor of Anatomy and Cellular Biology
Cell, Molecular & Developmental Biology  Neuroscience 
Olfactory system development, focusing on receptor cell genesis, maturation and senescence.

Johanna M. Seddon, M.D., Sc.M. Professor of Ophthalmology
Clinical Research
The causes and prevention of retina-macular degeneration.

Harry P. Selker, M.D., M.S.P.H. Professor of Medicine
Director of the Clinical Research Program
Clinical Research 
Clinical study design; mathematical models that predict clinical outcomes; decision support to improve diagnosis and triage.

Erik Selsing, Ph.D. Associate Professor of Pathology,
Director of the Genetics Program
Genetics  Immunology 
Recombination, expression, and mutation of antibody genes; regulation of the antibody and T-cell receptor repertoire.

* Richard I. Shader, M.D. Professor of Pharmacology and Experimental Therapeutics
Director of the Pharmacology and Experimental Therapeutics Program
Pharmacology & Experimental Therapeutics 
Clinical pharmacology; drug metabolism and drug interactions.

* Louis Shuster, Ph.D. Professor Emeritus of Pharmacology and Experimental Therapeutics
Biochemistry  Neuroscience&nbs